Evidence of rebbachisaurids and titanosauriforms (Dinosauria: Sauropoda) from the Early Cretaceous of Tunisia

Isolated sauropod remains including vertebrae and a humerus from the Aïn El Guettar Formation (Albian, Early Cretaceous) of Tunisia are described. Vertebrae include a slightly procoelous anterior caudal vertebra, amphicoelous middle caudal vertebrae, and strongly procoelous distal caudal vertebrae. The humerus has an anteroposteriorly compressed shaft, robust deltopectoral crest restricted laterally and prominent condyles bounding a distinct distal fossa. The morphological characters present in the specimens suggests that isolated remains can be referred to at least two distinct sauropod taxa. The anterior caudal vertebra is referred to Rebbachisauridae, whereas remaining caudal vertebrae show titanosauriform and titanosaurian derived features (anteriorly placed neural arches and, in the posterior vertebrae, distincly procoelous centra); finally, the humerus may pertain to a somphospondylian titanosauriform, perhaps the same taxon represented by the middle and posterior caudal vertebrae. This study introduces some of the oldest titanosauriform remains from Northern Africa and provides additional data on the stratigraphic and geographic distribution of this clade during the Early Cretaceous.